The rate of oviposition was determined by recording egg clutches deposited daily. To make this measurement, a plastic pot with a diameter of 25cm X depth of 10 cm containing 1 kg of a saturated grain Diet of 50% wheat bran, 30% alfalfa meal, and 20% corn meal (Hogsette 1992) was placed in the center of the cage on a table at a height of 40 cm. Individuals oviposited in the flutes (2 X 3 mm.) of two corrugated cardboardrolls rolls (egg collecting units ) measuring 2.5 cm indiameter X 3.5 cm in length that were taped to the inside of the pot approximately 3 cm above the moist media. The cardboard rolls were replaced daily, and the numbers of egg clutches were recorded.