Resistance or travel cost reflects the ecological cost
of travel through a pixel. In general resistance
increases with the risk of dying in the pixel and the
energetic cost of travel through the pixel.
Resistance decreases with how much food, water,
and cover an individual might find in the pixel, or
the probability of finding a mate and breeding in
an area that includes the pixel. It is not necessarily
related to the speed of travel through the pixel. In
fact, animals may tend to move quickly through