This study was supported by grant under Higher Institutions Centre of Excellence (HICoE) (Grant code: COE0010). We are grateful to Professor Datuk Dr. Mohd Harun Abdullah, Vice Chancellor, Universiti Malaysia Sabah and Professor Dr. Saleem Mustafa, Director, Borneo Marine Research Institute, Universiti Malaysia Sabah, for their encouragement and support. We thank all the staff, research assistant and students of Borneo Marine Research Institute, Universiti Malaysia Sabah for their cooperation in the experiment.