The gene will confer resistance when expressed either in the homozygous or heterozygous
state, although the resistance is better in the homozygous form (Okogbenin et al. 2007). Rabbi et al. (2014) found the CMD2 gene locus on linkage group 16, with a peak at 69.12 cM, which was close to the previously reported locus (Akano et al. 2002). The association decreased on both sides of the peak due to recombination between the markers and the resistance gene.