The high percentage of Q. seminulum (6.9–16.2%), and the relatively coarse grain size of the sediment, indicates an environment characterized by medium- to high-energy levels, typical of tidal channels (Serandrei-Barbero et al., 2004; Figure 4). Moreover, the high percentage of C. granosum (19.7–25.7%) that tolerates salinity changes (Canali et al., 2007), and the relatively low percentage of A. beccarii (35–44.8%) is indicative of fresh water inputs