However, bryophytes can (and many do) roll their
leaves, as do some vascular plants, and they have several
other related options to reduce the exposed surface area.
These include curling and contorting the leaves, a
mechanism that creates small air spaces and presumably
decreases air movement across the leaf surface. Others
appress their leaves closely to the stem, protecting the
upper surface from exposure and overlapping leaves
sufficiently to protect even portions of the back surface of
the leaf from exposure. And, despite their lack of
specialized energy-storing organs (with some exceptions),
they do have life cycle options.