the margin of LLNP affects the species richness and community structure of dung beetle and whether the species richness of dung beetles inhabiting natural forest is significantly higher than those in land-use system (agroforestry systems or annual cultures). (ii) How significant the contribution of agroecosystems to the total (regional) species richness of dung beetles is. (iii) What habitat variables will be the best predictors for explaining differences in dung beetle communities between land-use type.