Water quality parameters were maintained within the range recommended by Vega and de la Cruz (1988) for
− 1 the species. Dissolved oxygen was 7.25±0.25 mg l , temperature was maintained at 28.5±0.5 °C, and pH was 8.25±0.05. NH4–N had a range between 0.013+
− 1
0.005 mg l .