Spatial layers used in this study came from the following sources: (1) Land cover, downloaded from the ESA Global Land Cover 2009 Pro- ject; (2) elevation, downloaded from WorldClim (Hijmans et al., 2005); (3) KBAs, provided by CEPF (Birdlife International et al., 2013); (4) protected areas, downloaded from the World Database on Protected Areas (IUCN and UNEP, 2014); and (5) the human footprint index, de- veloped by the Socioeconomic Data and Applications Center of Colum- bia University (Sanderson et al., 2002).