The seed is the beginning of life for most flowering plants. The bulk of knowledge we have about seed development and structure has come from the study of agronomic crops as the result of our need to better provide food for the masses. While there is no less emphasis today on the need to improve our understanding of seed development in agronomic crops, there is an increasing demand for knowledge about other types of seed as well. Beautiful flowers in the garden, along the roadside, and in our homes provide aesthetically pleasing surroundings for humans. In the quest to beautify our surroundings there are infinite possibilities for research related to understanding the mysteries of seed development in floral crops. With more than 250,000 species of flowering plants, humans have only scratched the surface of knowledge about these fascinating organisms. The information provided in this chapter is a very brief glimpse into the complex world of seed development and structure in floral crops.