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With trackerless/DHT torrents, peer IP addresses are stored in the DHT using the BitTorrent infohash as the key. Since all a tracker does, basically, is respond to put/get requests, this functionality corresponds exactly to the interface that a DHT (distributed hash table) provides: it allows you to look up and store IP addresses in the DHT by infohash.
So a "get" request would look up a BT infohash and return a set of IP addresses. A "put" stores an IP address for a given infohash. This corresponds to the "announce" request you would otherwise make to the tracker to receive a dictionary of peer IP addresses.