Based on 16S rDNA sequence similarity, strains PR1, PR5 and SS5 were similar to Burkholderia kururiensis (≥ 99 % sequence similarity), strains TS1 and SS2 were similar to Klebsiella sp. (100 % and 99 % sequence similarity, respectively), strains PS5, SR6 and SS6 were similar to Sphingomonas sp. (99 % sequence similarity), strain TR4 was similar to Novosphingobium sp. (99 % sequence similarity), and strain RR3 was similar to Novosphingobium nitrogenifigens (100 % sequence similarity).