Aanen & Eggleton
(2006) revealed that cultivation of Termitomyces by termites originated in the African rain forests as the main centre and migrated to other geographical regions like Asia and Madagascar. However, molecular studies have revealed no identical sequences among the Termitomyces between Africa and Asia (Frøslev et al. 2003), demonstrating their geographical difference as well as a possible independent evolution. The life cycles and intricacies of coevolution between termites and Termitomyces (asexual and sexual phase) is still being debated (Rouland-Lefevre et al. 2002, Nobre & Aanen 2012). Recently, Nobre & Aanen (2012) argued that the termites harvest asexual spores of fungus along with lignocellulolytic enzymes of fungal and bacterial origin through faeces and foraged plant material to build the comb (fungus garden), which is comparable to an external rumen.