Since geophytes do not photosynthesize during the autumn-winter period, starch hydrolysis is the only source of soluble sugars. Thus, the dynamics of sugar accumulation in blubs is generally follow those of starch degradation. During storage, the total sugar concentration in blubs is not influenced by differences in the surrounding temperature [11], and the amount of sucrose is always greater than that of reducing sugars [32]. The activities of sucrose-cleaving enzymes, sucrose synthase and the invertases remain at low levels [22].