richness was twice as high in PC compared with WT reefs (Fig. 4): 44 species per site in WT (68–152 species per reef), compared with 89 species per site in PC (157–215 species per reef). Octocoral richness was also 30% lower in WT than PC (10 vs. 15 genera per site, 25 vs. 35 genera per reef). Richness of hard corals was highest on Hannah Reef, where the surveys yielded a total of 215 species, whereas the highest octocoral richness was found at Hay Reef (42 genera per reef). In contrast, the mean richness of macroalgal genera was 30% higher in WT than PC, whereas the richness of fish species was similar in both regions (Table 1).