We used somewhat different criteria to identify
adult females that did not breed for the two species because
(i) C. hoffmanni reared young throughout the annual cycle,
whereas B. variegatus reared young over a considerably shorter
period (see below) and (ii) B. variegatus females were easier to
locate than C. hoffmanni females. As a result, we considered an
adult female B. variegatus to not have reproduced if it was
observed without young on at least five occasions from January
to June (in May 2012), with at least 1 week elapsing between
consecutive observations. For C. hoffmanni, we considered an
adult female to not have bred in a given year if she was not
observed caring for a juvenile at least once in each of the four
annual quarters (i.e. 3-month periods). Choloepus hoffmanni not
meeting this criterion, or that were observed for