Over sixty lineages of both monocot and eudicot angiosperms have evolved a remarkable solution to maximize photosynthesis efficiency under low CO2 levels, high temperatures and/or drought: C4 photosynthesis [1]. The evolution of this modified photosyn- thetic pathway represents a wonderful example of convergent evolution. While the changes necessary for the transition from C3 to C4 photosynthesis are numerous, the trait has a wide phylo- genetic distribution across angiosperms, with 19 different plant