The recent history of nymphalid systematics provides excellent examples of synergistic collaboration between molecular and
morphological approaches when combined data yield stronger supported and better resolved results at various taxonomic levels (other examples hereof include Wahlberg and Nylin, 2003;Simonsen et al., 2006; Simonsen et al., 2010; Brower et al., 2010;Penz et al., 2011). Perhaps no single problem illustrates this better than the systematic history of the genus Bia.