Changes natural forest to other land-use type has significantly affect on species richness (ANOVA: F(3,8) = 7.46, p < 0.05, Figure 1a) but not on the abundance of dung beetles (Figure 1b). Although species richness was decreased from natural forest to open cultivated area, this study only detected a significant reduction of species richness on open cultivated area. Species richness at the natural forest, disturb forest and cacao agroforestry were nearly similar (Figure 1b