METHODS
Study area
Fieldwork was conducted in and around a privately
owned organic shade-grown cocoa (Theobroma cacao)
farm, in north-eastern Costa Rica (10.32°N, 83.59°W;
Figure 1). The region possesses a wet and warm climate
featuring a rainy season from mid- or late-April to
January, which is briefly interrupted by a dry period in
AugustorSeptember(Holdridge1967, Janzen1983).The
study area occurs within an agricultural landscape that
contains five habitat types: (1) cocoa trees grown under
overstorey of native and non-native trees, (2) tropical
forest occurring in narrow (20 m) riparian buffers and
small patches, (3) cattle pastures with scattered trees,
(4) monocultures of cultivated crops, and (5) human
development (i.e. housing) that border the study area.We
used 2013 RapidEye satellite imagery (IntraSearch Inc.)
with 5-m resolution and five-bandmultispectral imagery
to visually digitize polygons representing these habitat
types using ArcGIS 10.1 (ESRI, Redlands, CA, USA).
METHODSStudy areaFieldwork was conducted in and around a privatelyowned organic shade-grown cocoa (Theobroma cacao)farm, in north-eastern Costa Rica (10.32°N, 83.59°W;Figure 1). The region possesses a wet and warm climatefeaturing a rainy season from mid- or late-April toJanuary, which is briefly interrupted by a dry period inAugustorSeptember(Holdridge1967, Janzen1983).Thestudy area occurs within an agricultural landscape thatcontains five habitat types: (1) cocoa trees grown underoverstorey of native and non-native trees, (2) tropicalforest occurring in narrow (20 m) riparian buffers andsmall patches, (3) cattle pastures with scattered trees,(4) monocultures of cultivated crops, and (5) humandevelopment (i.e. housing) that border the study area.Weused 2013 RapidEye satellite imagery (IntraSearch Inc.)with 5-m resolution and five-bandmultispectral imageryto visually digitize polygons representing these habitattypes using ArcGIS 10.1 (ESRI, Redlands, CA, USA).
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