Attributes varied widely among the 3 forest states. In antinvaded
forest, ant activity was extremely high (Fig. 1)
and was reflected in the densities of yellow crazy ants
(mean [SE] = 918.2/m2 [283.8]) extracted from litter.
No yellow crazy ants occurred in intact sites, and only 2
individuals were found at ghosted sites. Associated with
high ant densities at ant-invaded sites was the complete
absence of red crab burrows and their near absence at
ghosted sites (Fig. 1). Crab burrow densities were much
higher in intact sites. With no or few red crabs at the
ant-invaded and ghosted sites, leaf litter, the mainstay
of the crab’s diet, was 2–3 times more abundant than
in intact sites (Fig. 1). Densities of litter invertebrates
(excluding crazy ants) were highest in ghosted sites and
lowest in intact sites (Fig. 1), which was consistent with
differences in standing litter masses (averaging 267, 291,
and 354 invertebrates/kg litter at ant-invaded, intact, and
ghosted sites, respectively).