The leaf width range
presented by den Hartog (1970) for H. ovalis does not include the
width of the H. ovalis subspecies linearis (1.5mm wide). Additionally,
the leaf asymmetry of H. johnsonii and the Antigua samples
(Fig. 1) is a feature not reported for H. ovalis (den Hartog, 1970;
Kuo, 2007) but leaf asymmetry has been seen in some populations
of H. ovalis worldwide, including Egypt, Thailand, China, Guam, the
Philippines and Fiji (Short, pers. obs.).