Nevertheless, the decrease at T1 in was not related to the increase in proline in the 12/cultivar and 14/culti-var plants ( s= 0.318 [proline] − 6.288; r2= 0.62; P < 0.05). At this earlier period, other components such as glycinebetaine, carbohydrates, amino acids, and macronutrients could have contributed to reducing the osmotic potential (Munns et al., 1979; Morgan,1992; Navarro et al., 2003) in these plant combinations.