5 There is a large literature on polyculture systems and their achievement of feed efficiency
and waste circulation, including the largely experimental integrated multitrophic
aquaculture systems (IMTA). Some relevant papers for our discussion include: Li
(1987), Milstein (1992), Kestemont (1995), Edwards (1998), Lazard and Dabbadie
(2002), and Chopin et al. (2007).