Finally, we compared the empirical D. to the distribution generated by randomization and
considered results significant for p < 0.05 using a one-sided test
(i.e., a high observed D.).
We applied a correction for multiple tests,
controlling for false discovery rate by setting a threshold of 0.05 (61
total tests: 13 species for which we performed one habitat pair
comparison plus 16 species for which we conducted three habitat
pair comparisons; Benjamini and Hochberg, 1995).