Macrofauna were described using abundance (A), biomass
(B), number of species (S) and Shannon–Wiener diversity index (H′). Differences in these univariate measures
between plots were tested using a one-way analysis of
variance (ANOVA). The molluscan biomass data were
not normally distributed so a non-parametric Kruskal–
Wallis test was used instead. A Mann–Whitney U test was
used to test for significant differences between the transects and time of collection (at spring or neap tides).