Area reduction is likely to affect dung beetles indirectly by reduction in food availability as the complete extinction of large mammals and the drastic decline
in bird populations in small forest fragments imply a lesser amount of excrement and carcasses to be used as resources by dung beetles.
As dung beetle body length is positively correlated with the amount of dung consumed and area requirements (Halffter and Matthews, 1966; Larsen et al., 2008), larger bodied beetles are more vulnerable to habitat fragmentation