Cellular fatty acid analysis of cells cultivated on ISP 2 agar medium for 4 days at 30 oC was performed by Chromatography (GLC) according to the instructions of the Microbial Identification System (MIDI) Sherlock version 6.0 [18] with the RTSBA6 MIDI database (Faculty of Science, King Mongkut' Institute of Technology Ladkrabang).