Plant tissue culture plays an important role in the production of agricultural and ornamental plants and in the manipulation of plants for improved agronomic performance. In vitro culture of plant cells and tissue has attracted considerable interest over recent years because it provides the means to study plant physiological and genetic processes in addition to offering the potential to assist in the breeding of improved cultivars by increasing genetic variability. Regenerated plants are expected to have the same genotype as the donor plant; however, in some cases somaclonal variants have been found among regenerated plants (Karp et al., 1987).