The study area (Fig. 1) covers three important Bolivian ecoregions: the Andean Piedmont, which includes the upper Isiboro-Secure Basin of the Bolivian Amazon, with altitudes of 221–260 m; the Inter-Andean Valleys, which includes the Río Grande Basin, with altitudes of 1505–4286 m; and the High-Andes ecoregion, with altitudes of 3719–4449 m.