Toes slender, not webbed, nor fringed; toes lengths I < II < V < III < IV; toe tips slightly pointed; subarticular tubercles large,rounded; sole of foot with distinct, approximately aligned tubercles; outer metatarsal tubercle very small, elliptical, flat; inner metatarsal tubercle large, elliptical, slightly elevated; sole of tarsus with few tubercles; inner tarsal fold distinct, approximately equal to the length of the tarsus.