In this regard, ethyl acetate has been shown to have an antifungal effect and might contribute to the biocontrol activity of W. anomalus during airtight storage of grain (Fredlund et al., 2004; Druvefors et al., 2005) while more recently Hua et al. (2014) stated that the biocontrol ability of W. anomalus can be attributed to the production of 2- phenylethanol, which affects spore germination, growth, toxin production, and gene expression in Aspergillus flavus.