We included only one species of Pachites in this study.
Based on a morphological analysis, Linder and Kurzweil (1994)
and Kurzweil et al. (1995) suggested that Pachites might be
a paraphyletic assemblage relative to Satyrium:(P. bodkinii
(P. appressa, Satyrium)). Paraphyly is a result of the many
differences in the column structure between the two species
of Pachites and the fact that some of the column features of
P. appressa resemble more those of Satyrium. Furthermore, an
elongated column was considered a synapomorphy for the sub-
tribe and thus not evidence for monophyly of Pachites, whereas
the subactinomorphic flower were seen as primitive and thus
also not as evidence. However, the new phylogenetic position of
Pachites suggests that both these features might be derived.
Nevertheless, considering the extent of morphological conver-gence in this clade, this should be tested in further molecular
analyses, which will depend on collection of the second species
of this rare and enigmatic genus.