In this study, we investigated the species boundaries between the sibling species pair A.
ocellaris and A. percula (Subgenus Actinicola) and three closely related species of the subgenus Phalerebus(A. akallopisos, A. perideraion, A. sandaracinos) by phylogenetic analysis of mitochondrial cytochrome band control region sequences.
These two subgenera show strong differences in their patterns of species boundaries.