External skin color (three measurements at three equidistant points on the equatorial region of each individual fruit) was measured on 15 fruit from each replicate using a color meter and recording CIE L*, a*, and b* values. Negative a* values indicate green and positive a* values red color. Higher positive b* values indicate a more yellow skin color and negative b* blue color. These values were then used to calculate hue angle, where 0◦ = red–purple; 90◦ = yellow;
180◦ = bluish green; and 270◦ = blue (McGuire, 1992), and Chroma, which indicates the intensity or color saturation.