B. anynana, like other butterflies, has two types of scales, cover and ground, which alternate within a row with cover scales partially covering the ground scales and the point where both scales attach to the wing membrane (Fig. 1 H and I and Fig. S1)
(28). Both cover and ground scales contain a lower lamina with a continuous smooth surface below a region composed of lon- gitudinal ridges and crossribs, collectively referred to as the upper lamina and connected to the lower lamina via pillars called trabeculae (Fig. 1J and Fig. S1) (6). scales