The exine is usually reticulate, but also foveolate, rugulate, perforate and psilate exine occurs in the subtribe. The
thickness of the exine is only 0.3 pm in tetrad grains of Randia urmata and in
the pantoporate monad grains of R. ruizianu, whereas the thickest exine occurs in
Posoqueria (2.8 pm) and Eucliniu (2.9 pm). Tectal excrescenses are generally absent.
However, three genera with pollen in tetrads, namely Euclinia, Gardenia and Oligoco-
don, show this feature. Contrary to the subtribe most genera are fairly stenopalynous,
but in the neotropical genera AIibertia and Rundia several different types of pollen are
encountered.