Xiu and Chai (2011) found that the modeled new production and the detrital nitrogen export in the cyclonic eddies were 28% and 41% higher than the SCS basin-averaged values, while in antic- yclonic eddies, those values were about 32% and 31% lower than the SCS basin mean. However, in sub-mesoscale domains of the anticyclonic eddies, Zhou et al. (2013) found enhanced Chl a and phytoplankton abundances stimulated by the upwelling at the eddy edge, as well as increased particle fluxes in the eddy core by horizontal advection