of Uca species on the same shore has been attributed to difference in shore height,
sediment type and/or salinity (e.g., Ono, 1962; Miller and Maurer, 1973; Icely and
Jones, 1978). Studies by Ono (1965) and Icely and Jones (1978) also showed that the
feeding appendages and gut structure of individual Uca species are adapted to feed
on a particular type of sediment. An examination of the buccal structures of the two