The authors would like to thank: Barbara East and Bryan Neves for greatly assisting the collection of field samples; Bill Trampe for use of facilities and access to study calves; and Dr Elizabeth Frazer for initiating the study and continued support. Funding for this research came from CSU College of Veterinary Medicine and Biomedical Sciences College Research Council and Integrated Livestock Management producer donations. Publication funding was generously provided by the Colorado State University Libraries Open Access Research and Scholarship Fund