The Hexapoda represent a monophyletic lineage within the Pancrustacea [1–3]. This lineage includes the insects and a number of apterygote groups such as Collembola, Diplura, Protura and Archaeognatha. Recent phylogenetic analysis has shown that Hexapoda represent one of six terrestrialization events within the Ecdysozoa [4]. According to this reconstruction, Hexapoda derive from a crustacean ancestor. This is supported by a study from Von Reumont et al., which suggests that hexapods most likely derive from a single terrestrialization event of an originally crustacean lineage