The soil mass, TC, TN, and TOC amounts excavated and deposited into burrow mimics, and net transport to the surface (that is, excavation minus deposition) per unit area were estimated, with these values being the highest in mudflats and the
lowest in Spartina marshes (Table 2). The soil mass per unit volume was the highest in mudflats and the lowest in Spartina marshes. The total C and N per unit volume were the highest in Phragmites