While abiotic factors like climate and topography limit species' distributions at coarse scales (Soberón, 2007), incorporating land cover and vegetation indices has improved the predictive power of niche models significantly (Waltari et al., 2014; Zimmermann et al., 2007).That said, however, ENMs based solely on land cover variables will not identify areas that were suitable in previous times (i.e., before land use change).