Raw forward and reverse sequences for each sample were assembled, ambiguous bases were corrected, and consensus sequences were edited using Sequencher 4.1 (Gene Codes Corporation, Ann Arbor, Michigan, USA). Consensus sequences for ITS and matK were manually aligned in Se-Al 2.0a11 (Rambaut, 2000⇓). All regions of ambiguous alignment within the ITS regions were excluded and gaps were treated as missing data.