morphology was examined using light microscopy
(E600; Nikon) and transmission electron microscopy (CM-
20; Philips). Flagellation was investigated using cells from
an exponentially growing culture. For this purpose, cells
were negatively stained with 1% (w/v) phosphotungstic
acid and the grids were examined after being air-dried.
Gliding motility was investigated as described by Bowman
(2000). The Gram reaction was investigated using the
bioMe´rieux Gram-stain kit, according to the manufacturer’s
instructions.