The four new species described in t

The four new species described in the present paper
may constitute a monophyletic group with such species as
Nemophora pruinosa Hirowatari, 2005 from the Ryukyus,
Japan, and N. umbripennis Stringer, 1930 from the main
islands of Japan. These species shares the following characteristics:
(1) the valvae are ventrally separated; (2) the
sacculus is basally somewhat swollen with dense hairs;
(3) the apical portion of the aedeagus bears a weak lamellate
keel dorsally, and (4) the manica has a patch of minute
spines dorsally. If these species comprise a monophyletic
group, the following scenario of diversification of this